After two years under Jim Mora, UCLA has become one of the hottest up-and-coming programs in the country. But the Bruins nonetheless have a major staffing question to answer after defensive coordinator Lou Spanos was named the Tennessee Titans' linebackers coach Saturday.

The Titans announced the hire on their official Twitter feed.

An NFL coaching veteran who spent 16 seasons with the Pittsburgh Steelers and Washington Redskins, Spanos signed on with the Bruins shortly after Mora's hire in late 2011. Under Spanos, UCLA improved from 87th in FBS yards-per-play allowed to 66th in 2012, then leapt all the way to 25th in 2013.

Many of UCLa's individual talents thrived under Spanos, most notably All-American linebacker Anthony Barr.