The 49ers placed fullback Bruce Miller on season-ending injured reserve and signed defensive lineman/fullback Will Tukuafu for the 53-man roster.

Tukuafu played 90 snaps of offense last season as a short-yardage blocker. But in the offseason program and training camp, he concentrated solely on the defensive side.

Tukuafu (6 foot 4, 293 pounds) did not play any offense during the four-game exhibition season while seeing 115 snaps of defense.

He was on the 49ers’ 53-man roster to open the season but was waived/injured on Sept. 10 with a knee injury. His injury settlement, which made allowed him to return to the 49ers later in the season, consisted of one game check.